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Cho KO, Kimura T, Ochiai K, et al. 
Gizzard adenocarcinoma in an aged Humboldt penguin (Spheniscus humboldti). [Journal Article]
Avian Pathol 1998 Feb; 27(1):100-2.
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Yonemaru K, Sakai H, Asaoka Y, et al. 
Proventricular adenocarcinoma in a Humboldt penguin (Spheniscus humboldti) and a great horned owl (Bubo virginianus); identification of origin by mucin histochemistry. [Case Reports, Journal Article, Research Support, Non-U.S. Gov't ]
Avian Pathol 2004 Feb; 33(1):77-81.
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Sivak J, Howland HC, McGill-Harelstad P 
Vision of the Humboldt penguin (Spheniscus humboldti) in air and water. [Journal Article]
Proc R Soc Lond B Biol Sci 1987 Jan 22; 229(1257):467-72.
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Hoogstraal H, Wassef HY, Hays C, et al. 
Ornithodoros (Alectorobius) spheniscus n. sp. [Acarina: Ixodoidea: Argasidae: Ornithodoros (Alectorobius) capensis group], a tick parasite of the Humboldt penguin in Peru. [Journal Article]
J Parasitol 1985 Oct; 71(5):635-44.
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Kikkawa EF, Tsuda TT, Naruse TK, et al. 
Analysis of the sequence variations in the Mhc DRB1-like gene of the endangered Humboldt penguin (Spheniscus humboldti). [Journal Article]
Immunogenetics 2005 Apr; 57(1-2):99-107.
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Rambaud YF, Flach EJ, Freeman KP 
Malignant melanoma in a Humboldt penguin (Spheniscus humboldti). [Case Reports, Journal Article]
Vet Rec 2003 Aug 16; 153(7):217-8.
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Nakanishi H, Hosoda S, Takahashi Y, et al. 
Alteration of tumour cell arrangement related to connective tissue stroma in metastatic brain tumours. Histological and immunohistochemical studies of 68 autopsy cases. [Journal Article]
Virchows Arch A Pathol Anat Histopathol 1989; 414(6):485-95.
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Bollmer JL, Vargas FH, Parker PG 
Low MHC variation in the endangered Galápagos penguin (Spheniscus mendiculus). [JOURNAL ARTICLE]
Immunogenetics 2007 Apr 25.
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Sano Y, Aoki M, Takahshi H, et al. 
The first record of Dirofilaria immitis infection in a Humboldt penguin, Spheniscus humboldti. [Case Reports, Journal Article]
J Parasitol 2005 Oct; 91(5):1235-7.
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